PARIS — King Richard I, the 12th century warrior whose bravery during the Third Crusade gained him the moniker Lionheart, ended up with a heart full of daisies, as well as myrtle, mint, and frankincense.

Those were among the finding of a French study, announced Thursday, which analyzed the embalmed heart more than 810 years after the king died.

The biomedical analysis also uncovered less flowery and spicy elements like creosote, mercury, and perhaps lime in the heart, which has been in the western French city of Rouen since his death in 1199.

Despite the embalming ingredients, the heart turned to powder long ago, doubtless because the lead box cradling it wasn't airtight.

The study's leader, Philippe Charlier, suggests the flowers and spices were to give the king the "odor of sanctity."